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on the relations of man to the lower animals-第5部分
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cavity of different races of mankind is far greater; absolutely; than
that between the lowest Man and the highest Ape; while; relatively; it
is about the same。 For the largest human skull measured by Morton
contained 114 cubic inches; that is to say; had very nearly double the
capacity of the smallest; while its absolute preponderance; of 52 cubic
inchesis far greater than that by which the lowest adult male human
cranium surpasses the largest of the Gorillas (62 … 34…1/2 = 27…1/2)。
Secondly; the adult crania of Gorillas which have as yet been measured
differ among themselves by nearly one…third; the maximum capacity being
34。5 cubic inches; the minimum 24 cubic inches; and; thirdly; after
making all due allowance for difference of size; the cranial capacities
of some of the lower Apes fall nearly as much; relatively; below those
of the higher Apes as the latter fall below Man。
Thus; even in the important matter of cranial capacity; Men differ more
widely from one another than they do from the Apes; while the lowest
Apes differ as much; in proportion; from the highest; as the latter
does from Man。 The last proposition is still better illustrated by the
study of the modifications which other parts of the cranium undergo in
the Simian series。
It is the large proportional size of the facial bones and the great
projection of the jaws which confers upon the Gorilla's skull its small
facial angle and brutal character。
FIG。 16。Sections of the skulls of Man and various Apes; drawn so as to
give the cerebral cavity the same length in each case; thereby
displaying the varying proportions of the facial bones。 The line 'b'
indicates the plane of the tentorium; which separates the cerebrum from
the cerebellum; 'd'; the axis of the occipital outlet of the skull。 The
extent of cerebral cavity behind 'c'; which is a perpendicular erected
on 'b' at the point where the tentorium is attached posteriorly;
indicates the degree to which the cerebrum overlaps the cerebellumthe
space occupied by which is roughly indicated by the dark shading。 In
comparing these diagrams; it must be recollected; that figures on so
small a scale as these simply exemplify the statements in the text; the
proof of which is to be found in the objects themselves。
But if we consider the proportional size of the facial bones to the
skull proper only; the little 'Chrysothrix' (Fig。 16) differs very
widely from the Gorilla; and; in the same way; as Man does; while the
Baboons ('Cynocephalus'; Fig。 16) exaggerate the gross proportions of
the muzzle of the great Anthropoid; so that its visage looks mild and
human by comparison with theirs。 The difference between the Gorilla
and the Baboon is even greater than it appears at first sight; for the
great facial mass of the former is largely due to a downward development
of the jaws; an essentially human character; superadded upon that
almost purely forward; essentially brutal; development of the same
parts which characterizes the Baboon; and yet more remarkably
distinguishes the Lemur。
Similarly; the occipital foramen of 'Mycetes' (Fig。 16); and still more
of the Lemurs; is situated completely in the posterior face of the
skull; or as much further back than that of the Gorilla; as that of the
Gorilla is further back than that of Man; while; as if to render patent
the futility of the attempt to base any broad classificatory
distinction on such a character; the same group of Platyrhine; or
American monkeys; to which the Mycetes belongs; contains the
Chrysothrix; whose occipital foramen is situated far more forward than
in any other ape; and nearly approaches the position it holds in Man。
Again; the Orang's skull is as devoid of excessively developed
supraciliary prominences as a Man's; though some varieties exhibit
great crests elsewhere (See pp。 231; 232); and in some of the Cebine
apes and in the 'Chrysothrix'; the cranium is as smooth and rounded as
that of Man himself。
What is true of these leading characteristics of the skull; holds good;
as may be imagined; of all minor features; so that for every constant
difference between the Gorilla's skull and the Man's; a similar
constant difference of the same order (that is to say; consisting in
excess or defect of the same quality) may be found between the
Gorilla's skull and that of some other ape。 So that; for the skull; no
less than for the skeleton in general; the proposition holds good; that
the differences between Man and the Gorilla are of smaller value than
those between the Gorilla and some other Apes。
In connection with the skull; I may speak of the teethorgans which
have a peculiar classificatory value; and whose resemblances and
differences of number; form; and succession; taken as a whole; are
usually regarded as more trustworthy indicators of affinity than any
others。
FIG。 17。Lateral views; of the same length; of the upper jaws of
various Primates。 'i'; incisors; 'c'; canines' 'pm'; premolars; 'm';
molars。 A line is drawn through the first molar of Man; 'Gorilla';
'Cynocephalus'; and 'Cebus'; and the grinding surface of the second
molar is shown in each; its anterior and internal angle being just
above the 'm' of 'm2'。
Man is provided with two sets of teethmilk teeth and permanent teeth。
The former consist of four incisors; or cutting teeth; two canines; or
eyeteeth; and four molars; or grinders; in each jawmaking twenty in
all。 The latter (Fig。 17) comprise four incisors; two canines; four
small grinders; called premolars or false molars; and six large
grinders; or true molars; in each jawmaking thirty…two in all。 The
internal incisors are larger than the external pair; in the upper jaw;
smaller than the external pair; in the lower jaw。 The crowns of the
upper molars exhibit four cusps; or blunt…pointed elevations; and a
ridge crosses the crown obliquely; from the inner; anterior cusp to the
outer; posterior cusp (Fig。 17 m2)。 The anterior lower molars have
five cusps; three external and two internal。 The premolars have two
cusps; one internal and one external; of which the outer is the higher。
In all these respects the dentition of the Gorilla may be described in
the same terms as that of Man; but in other matters it exhibits many
and important differences (Fig。 17)。
Thus the teeth of man constitute a regular and even serieswithout any
break and without any marked projection of one tooth above the level of
the rest; a peculiarity which; as Cuvier long ago showed; is shared by
no other mammal save oneas different a creature from man as can well
be imaginednamely; the long extinct 'Anoplotherium'。 The teeth of the
Gorilla; on the contrary; exhibit a break; or interval; termed the
'diastema'; in both jaws: in front of the eye…tooth; or between it and
the outer incisor; in the upper jaw; behind the eyetooth; or between it
and the front false molar; in the lower jaw。 Into this break in the
series; in each jaw; fits the canine of the opposite jaw; the size of
the eye…tooth in the Gorilla being so great that it projects; like a
tusk; far beyond the general level of the other teeth。 The roots of
the false molar teeth of the Gorilla; again; are more complex than in
Man; and the proportional size of the molars is different。 The Gorilla
has the crown of the hindmost grinder of the lower jaw more complex;
and the order of eruption of the permanent teeth is different; the
permanent canines making their appearance before the second and third
molars in Man; and after them in the Gorilla。
Thus; while the teeth of the Gorilla closely resemble those of Man in
number; kind; and in the general pattern of their crowns; they exhibit
marked differences from those of Man in secondary respects; such as
relative size; number of fangs; and order of appearance。
But; if the teeth of the Gorilla be compared with those of an Ape; no
further removed from it than a 'Cynocephalus'; or Baboon; it will be
found that differences and resemblances of the same order are easily
observable; but that many of the points in which the Gorilla resembles
Man are those in which it differs from the Baboon; while various
respects in which it differs from Man are exaggerated in the
'Cynocephalus'。 The number and the nature of the teeth remain the same
in the Baboon as in the Gorilla and in Man。 But the pattern of the
Baboon's upper molars is quite different from that described above
(Fig。 17); the canines are proportionally longer and more knife…like;
the anterior premolar in the lower jaw is specially modified; the
posterior molar of the lower jaw is still larger and more complex than
in the Gorilla。
Passing from the old…world Apes to those of the new world; we meet with
a change of much greater importance than any of these。 In such a genus
as 'Cebus'; for example (Fig。 17); it will be found that while in some
secondary points; such as the projection of the canines and the
diastema; the resemblance to the great ape is preserved; in other and
most important respects; the dentition is extremely different。 Instead
of 20 teeth in the milk set; there are 24: instead of 32 teeth in the
permanent set; there are 36; the false molars being increased from eight
to twelve。 And in form; the crowns of the molars are very unlike those
of the Gorilla; and differ far more widely from the human pattern。
The Marmosets; on the other hand; exhibit the same number of teeth as
Man and the Gorilla; but; notwithstanding this; their dentition is very
different; for they have four more false molars; like the other
American monkeysbut as they have four fewer true molars; the total
remains the same。 And passing from the American apes to the Lemurs;
the dentition becomes still more completely and essentially different
from that of the Gorilla。 The incisors begin to vary both in number
and in form。 The molars acquire; more and more; a many…pointed;
insectivorous character; and in one Genus; the Aye…Aye ('Cheiromys');
the canines disappear; and the teeth completely simulate those of a
Rodent (Fig。 17)。
Hence it is obvious that; greatly as the dentition of the highest Ape
differs from that of Man; it differs far more widely from that of the
lower and lowest Ape
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