yeast-第3部分

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would　go；　but　the　solid　parts　would　be　stopped　behind；　the　torula　would
be　stopped；　the　liquid　parts　of　the　yeast　would　go。　　And　then　he　took
another　vessel　containing　a　fermentable　solution　of　sugar；　and　he　put
one　inside　the　other；　and　in　this　way　you　see　the　fluid　parts　of　the
yeast　were　able　to　pass　through　with　the　utmost　ease　into　the　sugar；　but
the　solid　parts　could　not　get　through　at　all。　　And　he　judged　thus：　　if
the　fluid　parts　are　those　which　excite　fermentation；　then；　inasmuch　as
these　are　stopped；　the　sugar　will　not　ferment；　and　the　sugar　did　not
ferment；　showing　quite　clearly；　that　an　immediate　contact　with　the
solid；　living　torula　was　absolutely　necessary　to　excite　this　process　of
splitting　up　of　the　sugar。　　This　experiment　was　quite　conclusive　as　to
this　particular　point；　and　has　had　very　great　fruits　in　other
directions。

Well；　then；　the　yeast　plant　being　essential　to　the　production　of
fermentation；　where　does　the　yeast　plant　come　from？　　Here；　again；　was
another　great　problem　opened　up；　for；　as　I　said　at　starting；　you　have；
under　ordinary　circumstances　in　warm　weather；　merely　to　expose　some
fluid　containing　a　solution　of　sugar；　or　any　form　of　syrup　or　vegetable
juice　to　the　air；　in　order；　after　a　comparatively　short　time；　to　see
all　these　phenomena　of　fermentation。　　Of　course　the　first　obvious
suggestion　is；　that　the　torula　has　been　generated　within　the　fluid。　　In
fact；　it　seems　at　first　quite　absurd　to　entertain　any　other　conviction；
but　that　belief　would　most　assuredly　be　an　erroneous　one。

Towards　the　beginning　of　this　century；　in　the　vigorous　times　of　the　old
French　wars；　there　was　a　Monsieur　Appert；　who　had　his　attention
directed　to　the　preservation　of　things　that　ordinarily　perish；　such　as
meats　and　vegetables；　and　in　fact　he　laid　the　foundation　of　our　modern
method　of　preserving　meats；　and　he　found　that　if　he　boiled　any　of　these
substances　and　then　tied　them　so　as　to　exclude　the　air；　that　they　would
be　preserved　for　any　time。　　He　tried　these　experiments；　particularly
with　the　must　of　wine　and　with　the　wort　of　beer；　and　he　found　that　if
the　wort　of　beer　had　been　carefully　boiled　and　was　stopped　in　such　a　way
that　the　air　could　not　get　at　it；　it　would　never　ferment。　　What　was　the
reason　of　this？　　That；　again；　became　the　subject　of　a　long　string　of
experiments；　with　this　ultimate　result；　that　if　you　take　precautions　to
prevent　any　solid　matters　from　getting　into　the　must　of　wine　or　the　wort
of　beer；　under　these　circumstancesthat　is　to　say；　if　the　fluid　has
been　boiled　and　placed　in　a　bottle；　and　if　you　stuff　the　neck　of　the
bottle　full　of　cotton　wool；　which　allows　the　air　to　go　through　and
stops　anything　of　a　solid　character　however　fine；　then　you　may　let　it
be　for　ten　years　and　it　will　not　ferment。　　But　if　you　take　that　plug
out　and　give　the　air　free　access；　then；　sooner　or　later　fermentation
will　set　up。　　And　there　is　no　doubt　whatever　that　fermentation　is
excited　only　by　the　presence　of　some　torula　or　other；　and　that　that
torula　proceeds　in　our　present　experience；　from　pre…existing　torulae。
These　little　bodies　are　excessively　light。　　You　can　easily　imagine　what
must　be　the　weight　of　little　particles；　but　slightly　heavier　than　water；
and　not　more　than　the　two…thousandth　or　perhaps　seven…thousandth　of　an
inch　in　diameter。　　They　are　capable　of　floating　about　and　dancing　like
motes　in　the　sunbeam；　they　are　carried　about　by　all　sorts　of　currents
of　air；　the　great　majority　of　them　perish；　but　one　or　two；　which　may
chance　to　enter　into　a　sugary　solution；　immediately　enter　into　active
life；　find　there　the　conditions　of　their　nourishment；　increase　and
multiply；　and　may　give　rise　to　any　quantity　whatever　of　this　substance
yeast。　　And；　whatever　may　be　true　or　not　be　true　about　this
〃spontaneous　generation；〃　as　it　is　called　in　regard　to　all　other　kinds
of　living　things；　it　is　perfectly　certain；　as　regards　yeast；　that　it
always　owes　its　origin　to　this　process　of　transportation　or　inoculation；
if　you　like　so　to　call　it；　from　some　other　living　yeast　organism；　and
so　far　as　yeast　is　concerned；　the　doctrine　of　spontaneous　generation　is
absolutely　out　of　court。　　And　not　only　so；　but　the　yeast　must　be　alive
in　order　to　exert　these　peculiar　properties。　　If　it　be　crushed；　if　it　be
heated　so　far　that　its　life　is　destroyed；　that　peculiar　power　of
fermentation　is　not　excited。　　Thus　we　have　come　to　this　conclusion；　as
the　result　of　our　inquiry；　that　the　fermentation　of　sugar；　the
splitting　of　the　sugar　into　alcohol　and　carbonic　acid；　glycerine；　and
succinic　acid；　is　the　result　of　nothing　but　the　vital　activity　of　this
little　fungus；　the　torula。

And　now　comes　the　further　exceedingly　difficult　inquiryhow　is　it　that
this　plant；　the　torula；　produces　this　singular　operation　of　the
splitting　up　of　the　sugar？　　Fabroni；　to　whom　I　referred　some　time　ago；
imagined　that　the　effervescence　of　fermentation　was　produced　in　just　the
same　way　as　the　effervescence　of　a　sedlitz　powder；　that　the　yeast　was　a
kind　of　acid；　and　that　the　sugar　was　a　combination　of　carbonic　acid　and
some　base　to　form　the　alcohol；　and　that　the　yeast　combined　with　this
substance；　and　set　free　the　carbonic　acid；　just　as　when　you　add
carbonate　of　soda　to　acid　you　turn　out　the　carbonic　acid。　　But　of　course
the　discovery　of　Lavoisier　that　the　carbonic　acid　and　the　alcohol　taken
together　are　very　nearly　equal　in　weight　to　the　sugar；　completely　upset
this　hypothesis。　　Another　view　was　therefore　taken　by　the　French
chemist；　Thenard；　and　it　is　still　held　by　a　very　eminent　chemist；　M。
Pasteur；　and　their　view　is　this；　that　the　yeast；　so　to　speak；　eats　a
little　of　the　sugar；　turns　a　little　of　it　to　its　own　purposes；　and　by
so　doing　gives　such　a　shape　to　the　sugar　that　the　rest　of　it　breaks　up
into　carbonic　acid　and　alcohol。

Well；　then；　there　is　a　third　hypothesis；　which　is　maintained　by　another
very　distinguished　chemist；　Liebig；　which　denies　either　of　the　other
two；　and　which　declares　that　the　particles　of　the　sugar　are；　as　it
were；　shaken　asunder　by　the　forces　at　work　in　the　yeast　plant。　　Now　I
am　not　going　to　take　you　into　these　refinements　of　chemical　theory；　I
cannot　for　a　moment　pretend　to　do　so；　but　I　may　put　the　case　before　you
by　an　analogy。　　Suppose　you　compare　the　sugar　to　a　card　house；　and
suppose　you　compare　the　yeast　to　a　child　coming　near　the　card　house；
then　Fabroni's　hypothesis　was　that　the　child　took　half　the　cards　away；
Thenard's　and　Pasteur's　hypothesis　is　that　the　child　pulls　out　the
bottom　card　and　thus　makes　it　tumble　to　pieces；　and　Liebig's　hypothesis
is　that　the　child　comes　by　and　shakes　the　table　and　tumbles　the　house
down。　　I　appeal　to　my　friend　here　（Professor　Roscoe）　whether　that　is　not
a　fair　statement　of　the　case。

Having　thus；　as　far　as　I　can；　discussed　the　general　state　of　the
question；　it　remains　only　that　I　should　speak　of　some　of　those
collateral　results　which　have　come　in　a　very　remarkable　way　out　of　the
investigation　of　yeast。　　I　told　you　that　it　was　very　early　observed　that
the　yeast　plant　consisted　of　a　bag　made　up　of　the　same　material　as　that
which　composes　wood；　and　of　an　interior　semifluid　mass　which　contains　a
substance；　identical　in　its　composition；　in　a　broad　sense；　with　that
which　constitutes　the　flesh　of　animals。　　Subsequently；　after　the
structure　of　the　yeast　plant　had　been　carefully　observed；　it　was
discovered　that　all　plants；　high　and　low；　are　made　up　of　separate　bags
or　〃cells；〃　as　they　are　called；　these　bags　or　cells　having　the
composition　of　the　pure　matter　of　wood；　having　the　same　composition；
broadly　speaking；　as　the　sac　of　the　yeast　plant；　and　having　in　their
interior　a　more　or　less　fluid　substance　containing　a　matter　of　the　same
nature　as　the　protein　substance　of　the　yeast　plant。　　And　therefore　this
remarkable　result　came　outthat　however　much　a　plant　may　differ　from　an
animal；　yet　that　the　essential　constituent　of　the　contents　of　these
various　cells　or　sacs　of　which　the　plant　is　made　up；　the　nitrogenous
protein　matter；　is　the　same　in　the　animal　as　in　the　plant。　　And　not　only
was　this　gradually　discovered；　but　it　was　found　that　these　semifluid
contents　of　the　plant　cell　had；　in　many　cases；　a　remarkable　power　of
contractility　quite　like　that　of　the　substance　of　animals。　And　about　24
or　25　years　ago；　namely；　about　the　year　1846；　to　the　best　of　my
recollection；　a　very　eminent　German　botanist；　Hugo　Von　Mohl；　conferred
upon　this　substance　which　is　found　in　the　interior　of　the　plant　cell；
and　which　is　identical　with　the　matter　found　in　the　inside　of　the　yeast
cell；　and　which　again　contains　an　animal　substance　similar　to　that　of
which　we　ourselves　are　made　uphe　conferred　upon　this　that　title　of
〃protoplasm；〃　which　has　brought　other　people　a　great　deal　of　trouble
since！　　I　beg　particularly　to　say　that；　because　I　find　many　people
suppose　that　I　was　the　inventor　of　that　term；　whereas　it　has　been　in
existence　for　at　least　twenty…five　years。　　And　then　other　observers；
taking　the　question　up；　came　to　this　astonishing　conclusion　（working
from　this　basis　of　the　yeast）；　that　the　differences　between　animals　and
plants　are　not　so　much　in　the　fundamental　substances　which　compose　them；
not　in　the　protoplasm；　but　in　the　manner　in　which　the　cells　of　which
their　bodies　are　built　up　have　become　modified。　　There　is　a　sense　in
which　it　is　trueand　the　analogy　was　pointed　out　very　many　years　ago
by　some　French　botanists　and　chemiststhere　is　a　sense　in　which　it　is
true　that　every　plant　is　substantially　an　enormous　aggregation　of
bodies　similar　to　yeast　cells；　each　having　to　a　certain　extent　its　own
independent　life。　　And　there　is　a　sense　in　which　it　is　also　perfectly
truealthough　it　would　be　impossible　for　me　to　give　the　statement　to
you　with　proper　qualifications　and　limitations　on　an　occasion　like
thisbut　there　is　also　a　sense　in　which　it　is　true　that　every　animal
body　is　made　up　of　an　aggregation　of　minute　particles　of　protoplasm；
comparable　each　of　them　to　the　individual　separate　yeast　plant。　　And
those　who　are　acquainted　with　the　history　of　the　wonderful　revolution
which　has　been　worked　in　our　whole　conception　of　these　matters　in　the
last　thirty　years；　will　bear　me　out　in　saying　that　the　first　germ　of
them；　to　a